RESEARCH

Nature is full of diversity. Understanding this natural diversity is important to not only answer how different organisms evolve, but also to improve future conservation and agriculture. Our research is organized into three overarching themes.

Crop Genomics & Domestication

We apply advanced genomic assemblies to understand the history and future of our food. By moving beyond simple reference genomes to haplotype-resolved pangenomes, we are unlocking the genetic potential of established crops and accelerating the domestication of new ones.

Potato domestication history timeline
The Potato Pan-genome
Unraveling the complex autotetraploid genome of potato to understand its history and modernize its breeding.
View Project
Aardaker field
Aardaker: A New Protein Crop
Pioneering the domestication of Lathyrus tuberosus, a nitrogen-fixing tuber crop for sustainable protein.
View Project
Apple somatic mutation tree diagram
Apple Somatic Mutations
Uncovering the genomic basis of somatic "sport" mutations that drive cultivar evolution in apple trees.
View Project

The Potato Pan-genome

Craig Dent Sergio Tusso Lisa Baus Ana Kurdadze

The Challenge: Potato is a genetically complex autotetraploid. Standard sequencing collapses its four chromosome copies into one, losing critical variation.

Our Work: We constructed a haplotype-resolved pan-genome of European potato. By resolving all four haplotypes simultaneously, we revealed extraordinarily high sequence diversity between haplotypes — yet surprisingly low haplotype diversity across the continent, the fingerprint of historical breeding bottlenecks stretching back to the crop's introduction to Europe in 1570 and the devastating losses of the Irish Famine.

Key Publications
The phased pan-genome of tetraploid European potato. Sun H, Tusso S, Dent CI, Goel M, Wijfjes RY, Baus LC, Dong X, Campoy JA, Kurdadze A, Walkemeier B, Sänger C, Huettel B, Hutten RCB, van Eck HJ, Dehmer KJ, Schneeberger K.
Nature (2025).
Tracing modern breeding introgressions in European potato. Craig I Dent, Lisa C Baus, Sergio Tusso, Klaus J Dehmer, Ronald C B Hutten, Herman J van Eck, Korbinian Schneeberger.
Theoretical and Applied Genetics (2026).
Timeline of potato domestication and breeding history from South American origin ~10,000 years ago through European introduction in 1570, the Irish Famine in the mid-19th century, and the start of modern breeding in 1880.
History of European potato. Domesticated ~10,000 years ago in South America and introduced to Europe around 1570, potato became a continental staple before P. infestans caused the Irish Famine and wiped out susceptible genotypes. Modern breeding has proceeded largely within the European gene pool since ~1880, with only a handful of wild introgressions — shaping the narrow haplotype diversity we observe today.

Aardaker: The Protein Potato

Ana Kurdadze

While nature offers thousands of edible plant species, global agriculture is built around a small set of widely cultivated staple crops. We are working to change that. By harnessing the power of modern genomics, we aim to unlock this untapped potential and revive forgotten crops through de novo domestication.

Meet the Aardaker (L. tuberosus). Native to Eurasia, this member of the Fabaceae family is a nitrogen-fixing powerhouse that produces protein-rich tubers. We are generating a chromosome-scale reference genome and conducting a holistic population genomics analysis to uncover the species' evolutionary origins, characterize its existing genetic architecture, and assess its adaptive potential.

We work directly with breeders and openly share our insights to ensure the Aardaker makes the leap from scientific curiosity to a resilient, stable crop. More on the applied work at Aardaia →

Aardaker field
Aardaker field
Selected tubers from a single plant
Selected tubers from single plant
Aardaker pollinator
Aardaker pollinator
Harvested tubers
Harvested tubers
The protein potato
The protein potato
Root nodules for nitrogen fixation
Root nodules for nitrogen fixation
Young aardaker plant
Young aardaker plant
Large aardaker plant in the greenhouse
Large aardaker plant in the greenhouse

Apple Somatic Evolution

The Mystery of "Sport" Mutations: Many fruit varieties, such as the dark-red 'RubyMac' apple, arise spontaneously as somatic mutations on a single branch of a tree. Understanding these "bud sports" is crucial for breeding improved cultivars with desirable traits.

Genomic Forensics: We sequenced the genome of a single apple tree to identify the specific mutations separating wild-type and mutant branches. We discovered that somatic evolution is driven not just by new mutations, but frequently by gene conversions (loss of heterozygosity) that are specific to individual cell layers in the meristem.

Key Publication
The identification and analysis of meristematic mutations within the apple tree that developed the RubyMac sport mutation. Sun H, Abeli P, Campoy JA, Rütjes T, Krause K, Jiao WB, Beaudry R, Schneeberger K.
BMC Plant Biology (2024).
Two-dimensional projection of the apple tree showing sampling points and meristem model with three cell layers (L1, L2, L3).
Sampling design and meristem model. Pink: mutant (red apple) scaffolds; green: wildtype. Circles show approximate regions selected for whole-genome sequencing. Inset: three-layered meristem model and corresponding leaf cell layers.

Meiotic Dynamics & Evolution

Genetic diversity is driven by the reshuffling of chromosomes during meiosis. We pioneer single-cell sequencing technologies to understand the controlled (recombination) and uncontrolled (mutations) events that drive genomic diversity, and finally build the basis for all the variation in life.

Experimental workflow for single-nucleus pollen sequencing
Single-Nucleus Pollen Sequencing
Reconstructing meiotic recombination events by sequencing individual pollen grains, bypassing F2 populations.
View Project
Violin plot showing estimated crossovers across F1 hybrids
Natural Variation in Recombination
Investigating why recombination landscapes differ between individuals and identifying the genetic modifiers involved.
View Project
Centromere mutation heatmap and HOR score plot
Centromere Evolution
Using long-read assemblies to understand the rapid evolution of centromeric repeats and their role in speciation.
View Project

Single-Nucleus Pollen Sequencing

Samija Amar Matthew T. Parker Jonas Freudigmann

A Versatile Platform for Haploid Genetics: We are pioneering single-nucleus sequencing technologies to study the haploid life cycle of plants. By isolating and sequencing individual pollen nuclei, we can treat each gamete as an independent "offspring." This allows us to perform high-resolution genetic mapping without the need for time-consuming crosses or large progeny populations.

Beyond Recombination: While our primary focus has been mapping crossover landscapes, this technology is broadly applicable. We develop methods to analyze gene expression dynamics during pollen development, characterize rare cell types, and investigate the transcriptomic consequences of haploid selection. Our group actively maintains the tools (e.g., coelsch) required to analyze these complex, sparse molecular datasets.

Key Publications
coelsch: Platform-agnostic single-cell analysis of meiotic recombination. Parker MT, Amar S, Freudigmann J, Walkemeier B, Dong X, Solier V, Marek M, Huettel B, Mercier R, Schneeberger K.
preprint (2026).
Scalable eQTL mapping using single-nucleus RNA-sequencing of recombined gametes from a small number of individuals. Parker MT, Amar S, Campoy JA, Krause K, Tusso S, Marek M, Huettel B, Schneeberger K.
PLOS Biology (2025).
Experimental workflow: F1 hybrids generated, pollen nuclei isolated for droplet- or plate-based single-cell sequencing, reads aligned to parental genomes to reconstruct recombinant haplotypes.
Experimental workflow for crossover detection in recombinant Arabidopsis thaliana pollen. F₁ hybrids are generated and pollen nuclei isolated for droplet- or plate-based single-cell sequencing.

Natural Variation in Recombination

Matthew T. Parker Raúl Wijfjes Jonas Freudigmann

Population-Scale Mapping: We investigate how genetic diversity shapes the landscape of meiotic recombination. By extending our single-cell framework to natural populations, we can identify the specific genetic modifiers — like trans-acting factors or local structural variants such as inversions — that control where and how frequently crossovers occur.

Genetics of the Haploid Phase: Beyond recombination, we map the regulatory networks of the gamete itself. Using single-nucleus eQTL mapping, we uncover how cis- and trans-regulatory variation drives gene expression in pollen. For example, we recently identified DUO3 as a master regulator of sperm cell development by linking haplotype inheritance to transcriptomes in single nuclei.

Interspecific variation in recombination: Species of the Brassicaceae family vary in genome structure and ecological characteristics, while their core meiotic genes are well-conserved. We aim to identify how this variation drives differences in recombination landscape between species and the degree to which specific factors (mating system, ploidy, chromosome number, translocations) contribute.

Key Publications
coelsch: Platform-agnostic single-cell analysis of meiotic recombination. Parker MT, Amar S, Freudigmann J, Walkemeier B, Dong X, Solier V, Marek M, Huettel B, Mercier R, Schneeberger K.
preprint (2026).
Scalable eQTL mapping using single-nucleus RNA-sequencing of recombined gametes from a small number of individuals. Parker MT, Amar S, Campoy JA, Krause K, Tusso S, Marek M, Huettel B, Schneeberger K.
PLOS Biology (2025).
Violin plot showing estimated number of crossovers per male meiosis for 34 F1 hybrids across 20 parent 2 genotypes, comparing Col0 and Cvi0 parents.
Estimated crossover counts per male meiosis across 34 F₁ hybrids. Violins trimmed to observed data range; blue = Col0 parent 1, orange = Cvi0 parent 1.

Centromere Evolution

Xiao Dong

Deciphering Centromere Dynamics: Centromeres are essential for cell division but evolve rapidly. We use replicated, error-free genome assemblies to track the "birth" of mutations in these repetitive regions. Our work reveals a unique mutational spectrum dominated by non-allelic gene conversions and small, structure-preserving indels.

From Chaos to Order: We model how these fundamental processes shape genome architecture. Our simulations demonstrate that simple, small-scale mutations are sufficient to drive the self-organization of complex, megabase-scale Higher-Order Repeat (HOR) structures, providing a unified theory for centromere evolution.

Key Publication
The mutational dynamics of the Arabidopsis centromeres. Dong X, Jiao WB, Campoy JA, Rabanal F, Ton J, Smith LM, Weigel D, Schneeberger K.
bioRxiv (2025).
Mutation patterns in the centromere of chromosome 4, showing point mutations (orange), insertions (red), and deletions (blue) alongside HOR score plot and sequence identity heatmap.
Mutation map and repeat structure of Arabidopsis Chr4 centromere. Circles: point mutations (orange), insertions (red), deletions (blue). Heatmap shows pairwise sequence identity across 10 kb windows.

Methods & Computational Tools

Modern biology is data-driven. We develop the algorithms required to analyze complex structural variations and graph genomes, while simultaneously establishing wet-lab protocols for next-generation sequencing applications.

Pan-genome gene composition across 69 Arabidopsis accessions
Pan-genomics
Moving beyond the linear reference genome to capture the full spectrum of genetic diversity within and between species.
View Project
Visualizing structural rearrangements using plotsr across 6 human genomes
Genomics Tools
Developing standard software like SyRI and Plotsr for structural variant analysis and visualization.
View Project
Bandage genome assembly graph visualization
Sequencing Methods & Genome Assemblies
Developing robust pipelines for difficult samples, single-cell applications, and long-read technologies.
View Project

Pan-genomics

Raúl Wijfjes Craig Dent Leon Rauschning Lisa Baus

Reference Bias: A single linear reference genome captures only a fraction of a species' genetic diversity. It introduces reference bias, causing us to miss genes and structural variants present in the population but absent from the reference.

Aggregating Diversity: Our group develops pan-genomic approaches to comprehensively characterize variation across plant populations and species. By combining long-read sequencing with advanced phasing methods, we construct haplotype-resolved assemblies that reveal the full spectrum of genetic variation.

Evolution of Gene Synteny: Conservation or loss of gene synteny between species can affect gene expression, and subsequently phenotype. Using a panel of diverse Brassicaceae genomes, we investigate conservation of gene synteny across 20 million years of evolution and its implications for gene function.

From chromosome-level pan-genomes of Arabidopsis to phased assemblies of tetraploid potato and comparative resources across the Brassicaceae, our work connects natural variation to phenotypic diversity.

Key Publications
The phased pan-genome of tetraploid European potato. Sun H, Tusso S, Dent CI, Goel M, Wijfjes RY, Baus LC, Dong X, Campoy JA, Kurdadze A, et al., Schneeberger K.
Nature (2025).
A pan-genome of 69 Arabidopsis thaliana accessions reveals a conserved genome structure throughout the global species range. Lian Q, Huettel B, Walkemeier B, Mayjonade B, Lopez-Roques C, Gil L, Roux F, Schneeberger K & Mercier R.
Nature Genetics (2024).
Annotated protein-coding gene composition across 69 Arabidopsis accessions, showing core (red), softcore (blue), dispensable (green), and private (purple) genes per accession.
Gene composition across 69 A. thaliana accessions. Red: core genes shared across all accessions; blue: softcore; green: dispensable; purple: private genes unique to single accessions.

Software Infrastructure

Manish Goel Matthew T. Parker Leon Rauschning

We develop methods that leverage genomic differences from SNPs to large chromosomal rearrangements. Our most popular tools include:

  • SyRI: Synteny and Rearrangement Identifier — finds inversions, translocations, and duplications between whole-genome assemblies.
  • plotsr: Visualizes structural similarities and rearrangements between multiple genomes in a single, publication-ready figure.
  • coelsch: A set of tools for identifying recombination events at the single nucleus/cell level.
Key Publications
SyRI: finding genomic rearrangements and local sequence differences from whole-genome assemblies. Goel M, Sun H, Jiao WB, Schneeberger K.
Genome Biology (2019).
plotsr: visualizing structural similarities and rearrangements between multiple genomes. Goel M, Schneeberger K.
Bioinformatics (2022).
coelsch: Platform-agnostic single-cell analysis of meiotic recombination. Parker MT, Amar S, Freudigmann J, Walkemeier B, Dong X, Solier V, Marek M, Huettel B, Mercier R, Schneeberger K.
preprint (2026).
Scalable eQTL mapping using single-nucleus RNA-sequencing of recombined gametes from a small number of individuals. Parker MT, Amar S, Campoy JA, Krause K, Tusso S, Marek M, Huettel B, Schneeberger K.
PLOS Biology (2025).
Visualizing structural rearrangements using plotsr across 10 chromosomes from 6 human genomes, with gene, SNP density, and centromere tracks.
plotsr output visualizing syntenic regions and structural rearrangements across 10 chromosomes from 6 human genomes. Optional annotation tracks show genes (with CDS and transcribed regions), SNP density, and centromeric regions — generated without further modifications.

Advanced Sequencing Methods

Lisa Baus Samija Amar Xiao Dong Sergio Tusso Craig Dent

A core part of our work is the continuous development and optimization of sequencing protocols and analytical pipelines, ensuring our datasets push the boundaries of what genome assembly can resolve.

Long- and ultra-long-read sequencing: We work to improve nanopore-based sequencing workflows for generating longer, more contiguous reads — a critical step for assembling the highly repetitive regions, centromeres, and structural variants that short-read methods miss entirely.

Single-cell and single-nucleus methods: We develop and refine protocols for single-cell and single-nucleus sequencing applications, from library preparation through to computational analysis. This includes both transcriptomic and genomic approaches tailored for sparse, low-input data.

Crucially, we do not limit these efforts to tractable model organisms. We actively scale and adapt our methods for complex crop genomes — including the highly polyploid potato and the emerging crop Lathyrus tuberosus (Aardaker) — where the genomic complexity demands purpose-built solutions rather than off-the-shelf pipelines.

All assemblies, annotations, and pipelines are shared openly to support the broader community in trait discovery, evolutionary genomics, and plant breeding.

Bandage visualization of a draft genome assembly graph
Example draft assembly graph visualized in Bandage, showing contig connectivity and assembly structure. Bandage (Wick et al.) is a tool for exploring genome assembly graphs; source available at github.com/rrwick/Bandage.
Looking for our software? SyRI, plotsr, coelsch and more on the dedicated Tools page.

Our Funding